Heroes and History

Physiology research has been an important element of our school for a century. The intersections of lung physiology and mechanics were first brought to clear fruition here in the early part of the twentieth century by the Drinker brothers, when Philip the engineer and Cecil the physician combined their talents to invent the iron lung. Around the middle of the century, Jim Whittenberger recognized the need to better understand lung biology before we could anticipate and interpret how environmental exposures threatened our health, and he was perhaps the first to include a program in physiology research in a department of environmental health. A crucial early recruitment of his was Jere Mead, who elucidated the mechanics of respiration through the middle and later parts of the century. In the late 1960s, Joe Brain began extending levels of lung biology analyses from organism and organs to cell and molecules. In the present, we are elucidating lung biology at ever more sophisticated levels of molecular detail, with unprecedented abilities to integrate this knowledge with cell, organ, organism, and population biology. The interdisciplinary nature of our earliest research, particularly the intersections of biology and engineering, continues to be a hallmark of our program. While excited about what we are now doing and planning, we are also proud of what our program has accomplished.

Where are they now? Below is a list of a few previous members of the program…


Jere Mead, M.D. (faculty)
In Memoriam

Mary Ellen Avery, M.D. (trainee)
In Memoriam
Children’s Hospital, Boston


Peter T. Macklem, M.D. (trainee)
In Memoriam
McGill University

Mary Ellen Wohl, M.D. (trainee)
In Memoriam
Professor of Pediatrics, Harvard Medical School


Everett Sinnett, Ph.D. (trainee)
Scientific Review Officer, Respiratory Sciences
National Institutes of Health

Julian Solway, M.D. (trainee)
Walter L. Palmer Distinguished Service Professor of Medicine
Chairman, Committee on Molecular Medicine
University of Chicago

Thomas Zeltner, M.D. (trainee)
Former State Secretary and Director
Federal Office of Public Health, Switzerland

Robert Banzett, Ph.D. (trainee, faculty)
Director of Dyspnea Lab, Beth Israel Deaconess Medical Center
Associate Professor of Medicine, Harvard Medical School

Stephen Loring, M.D. (trainee, faculty)
Associate Professor
Harvard Medical School
Beth Israel Deaconess Hospital


George Barnas, Ph.D. (trainee)
Scientific Review Officer
National Institutes of Health

Marc Hershenson, M.D. (trainee)
Frederick G.L. Huetwell Professor of Pediatrics
University of Michigan

Mara Ludwig, M.D. (trainee)
Professor of Medicine
McGill University

Michael B. Reid, Ph.D. (trainee, faculty)
Dean and Professor, Department of Applied Physiology and Kinesiology
University of Florida

Dimitrije Stamenovic, Ph.D. (trainee)
Associate Professor of Biomedical Engineering
College of Engineering, Boston University

Angeline E. Warner, D.V.M., Sc.D. (trainee, faculty)
Associate Professor and Associate Dean for Academic Affairs
Tufts Cummings School of Veterinary Medicine


Robert Clarke, Ph.D. (trainee)
Pulmatrix, Inc.

Claire M. Doerschuk, M.D. (faculty)
Professor of Medicine and Pathology, UNC at Chapel Hill School of Medicine
Vice-Chair of Research and Chief of the Division of Integrative Biology, Rainbow Babies Children’s Hospital

Ben Fabry, Ph.D. (trainee)
Director, Center for Medical Physics and Technology
Biophysics Group, Erlangen University, Germany

Charles W. Frevert, D.V.M., Sc.D. (trainee)
Research Associate Professor of Medicine
University of Washington

Geoff Maksym, Ph.D. (trainee)
Assistant Professor
School of Biomedical Engineering, Dalhousie University

Paul Moore, M.D. (trainee)
Assistant Professor of Pediatrics and Pharmacology
Vanderbilt University School of Medicine

Joseph Paulauskis, Ph.D. (faculty)
Chief Operating Officer

Steve Shea, Ph.D. (trainee)
Director, Center for Research on Occupational and Environmental Toxicology (CROET)
Oregon Health & Science University

Peggy Taylor, Ph.D. (trainee)
Vice President, Production and Development
Signet Laboratories

Andrew Binks, Ph.D. (Trainee)
Research Associate Professor
University of South Carolina School of Medicine
Greenville, SC

Karleyton Evans, MD (Trainee)
Laboratory for Neuroimaging of Anxiety and Respiratory Psychophysiology
Massachusetts General Hospital, Psychiatric Neuroimaging Division

Shakeeb Moosavi, Ph.D. (Trainee)
Senior Lecturer in Clinical Physiology
Department of Biological and Medical Sciences – Faculty of Health and Life Sciences
Oxford Brooks University


Ning Wang, Sc.D. (trainee, faculty)
Professor of Mechanical and Industrial Engineering
University of Illinois-Urbana

Steven An, Ph.D. (trainee)
Assistant Professor of Environmental Health Sciences
Johns Hopkins Bloomberg School of Public Health

Richard Johnston, Ph.D. (trainee)
Assistant Professor and Associate Director for Research
University of Texas Health Science Center at Houston

Andrea Vovk, Ph.D. (Trainee)
Clinical Research Consultant
University of Florida